The marginal wetlands that remain continue to be threatened by reclamation for farms and homes.

怎么样理解此处的remain continue？难道remain后面可以接动词吗？that remain不是做The marginal wetlands谓语吗？continue这里又是一个谓语？双谓语吗？

The marginal wetlands that remain continue to be threatened by reclamation for farms and homes.that remain是定语从句，谓语是continue。